Sunday, January 3, 2010

Jellyfish

Jellyfish (also jellies or sea jellies) are free-swimming members of the phylum Cnidaria. Jellyfish have several different morphologies that represent several different cnidarian classes including the Scyphozoa (over 200 species), Staurozoa (about 50 species), Cubozoa (about 20 species), and Hydrozoa (about 1000-1500 species that make jellyfish and many more that do not). The jellyfish in these groups are also called, respectively, scyphomedusae, stauromedusae, cubomedusae, and hydromedusae; medusa is another word for jellyfish, but it also refers specifically to the adult stage of the life cycle.
Jellyfish are found in every ocean, from the surface to the deep sea. Some hydrozoan jellyfish, or hydromedusae, are also found in fresh water and are less than half an inch in size. They are partially white and clear and do not sting. Many of the best-known jellyfish, such as Aurelia, are scyphomedusae. These are the large, often colorful, jellyfish that are common in coastal zones worldwide.
In its broadest sense, the term jellyfish also generally refers to members of the phylum Ctenophora. Although not closely related to cnidarian jellyfish, ctenophores are also free-swimming planktonic carnivores, are generally transparent or translucent, and exist in shallow to deep portions of all the world's oceans.

Life cycle
Most jellyfish undergo two distinct life history stages (body forms) during their life cycle. The first is the polypoid stage, when the animal takes the form of a small stalk with feeding tentacles; this polyp may be sessile, living on the bottom or on similar substrata such as floats or boat-bottoms, or it may be free-floating or attached to tiny bits of free-living plankton or rarely, fish. Polyps generally have a mouth surrounded by upward-facing tentacles like miniatures of the closely-related anthozoan polyps (sea anemones and corals), also of the phylum Cnidaria. Polyps may be solitary or colonial, and some bud asexually by various means, making more polyps. Most are very small, measured in millimeters.
In the second stage, the tiny polyps asexually produce jellyfish, each of which is also known as a medusa. Tiny jellyfish (usually only a millimeter or two across) swim away from the polyp and then grow and feed in the plankton. Medusae have a radially symmetric, umbrella-shaped body called a bell, which is usually supplied with marginal tentacles - fringe-like protrusions from the bell's border that capture prey. A few species of jellyfish do not have the polyp portion of the life cycle, but go from jellyfish to the next generation of jellyfish through direct development of fertilized eggs.
Jellyfish are dioecious; that is, they are either male or female. In most cases, both release sperm and eggs into the surrounding water, where the (unprotected) eggs are fertilized and mature into new organisms. In a few species, the sperm swim into the female's mouth fertilizing the eggs within the female's body. In moon jellies, the eggs lodge in pits on the oral arms, which form a temporary brood chamber.
After fertilization and initial growth, a larval form, called the planula, develops. The planula is a small larva covered with cilia. It settles onto a firm surface and develops into a polyp. The polyp is cup-shaped with tentacles surrounding a single orifice, resembling a tiny sea anemone.
After a growth interval, the polyp begins reproducing asexually by budding and, in the Scyphozoa, is called a segmenting polyp, or a scyphistoma. New scyphistomae may be produced by budding or new, immature jellies called ephyrae may be formed. A few jellyfish species can produce new medusae by budding directly from the medusan stage. Budding sites vary by species; from the tentacle bulbs, the manubrium (above the mouth), or the gonads of hydromedusae. A few of species of hydromedusae reproduce by fission (splitting in half.)
Other species of jellyfish are among the most common and important jellyfish predators, some of which specialize in jellies. Other predators include tuna, shark, swordfish, sea turtles and at least one species of Pacific salmon. Sea birds sometimes pick symbiotic crustaceans from the jellyfish bells near the sea's surface, inevitably feeding also on the jellyfish hosts of these amphipods or young crabs and shrimp.
Jellyfish lifespans typically range from a few hours (in the case of some very small hydromedusae) to several months. Life span and maximum size varies by species. One unusual species is reported to live as long as 30 years. Another species, Turritopsis dohrnii as T. nutricula, may be effectively immortal because of its ability to transform between medusa and polyp, thereby escaping death. Most large coastal jellyfish live 2 to 6 months, during which they grow from a millimeter or two to many centimeters in diameter. They feed continuously and grow to adult size fairly rapidly. After reaching adult size, jellyfish spawn daily if there is enough food. In most species, spawning is controlled by light, so the entire population spawns at about the same time of day, often at either dusk or dawn.

Toxicity to humans
Scyphozoan jellyfish stings are not generally deadly, though some species of the completely separate class Cubozoa (box jellyfish), such as the famous and especially toxic Irukandji, can be.
Jellyfish sting using microscopic cells called nematocysts, which are capsules full of poison expelled through a microscopic lance. Contact with a jellyfish tentacle can trigger millions of nematocysts to pierce the skin and inject venom.
Jellyfish sting prey and threatening humans using their nematocysts, but only some jellyfish species harm humans. Even beached and dying jellyfish can still sting when touched. Sting effects range from no effect to extreme pain to death.
Stings may cause anaphylaxis, which may result in death. Hence, victims should immediately get out of the water. Medical care may include administration of an antivenom.
The three goals of first aid for uncomplicated jellyfish stings are: prevent injury to rescuers, inactivate the nematocysts, and remove tentacles attached to the patient. Rescuers should wear barrier clothing, such as panty hose, wet suits or full-body sting-proof suits. Inactivating the nematocysts, or stinging cells, prevents further injection of venom.
 
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